瘤拟步行虫科
瘤拟步行虫科
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| Ironclad beetle,（Zopherus nodulosus haldemani），摄于美国德州 | |
科学分类 
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| 界： | 动物界 Animalia |
| 门： | 节肢动物门 Arthropoda |
| 纲： | 昆虫纲 Insecta |
| 目： | 鞘翅目 Coleoptera |
| 亚目： | 多食亚目 Polyphaga |
| 下目： | 扁虫下目 Cucujiformia |
| 总科： | 拟步总科 Tenebrionoidea |
| 科： | 瘤拟步行虫科 Zopheridae Solier, 1834 |
| 亚科 | |
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Colydiinae - cylindrical bark beetles | |
瘤拟步行虫科（学名：Zopheridae）是属于鞘翅目拟步总科的昆虫。近年来，有其他两个科被划分到本科之中，使得本科的成员数量有了相当的成长；这两个科分别是拟吉丁虫科（Monommatidae）以及细坚虫科（Colydiidae）。某些学者接受本科划分了6个亚科的系统，然而有些则将细坚虫科从本科中剔除为一独立科。本科有超过100个属群需要被重新定义；据估计在世界各地有著数百个种类。这个科没有固定使用的英文俗名，不过多数亚科都有各自传统的英文俗名，例如 ironclad beetles、cylindrical bark beetles等。至2018年为止，约有116个物种已被描述。 [1] [2]
系统分类学
目前所使用的瘤拟步行虫科级分类元定义主要来自史利宾斯基和劳伦斯于 1999 年 基于成虫和幼生期形态所做的拟步行虫总科系统发育分析[3]，近年的分子系统发育分析研究则显示本科可能并非单系群[4][5][6]。
本科包括以下属：
- Ablabus Broun, 1880
- Acolobicus Sharp, 1894
- Acolophoides Slipinski & Lawrence, 1997
- Acolophus Sharp, 1885
- Acostonotus Slipinski & Lawrence, 1997
- Acropis Burmeister, 1840
- Afrorthocerus Pope, 1959
- Allobitoma Broun, 1921
- Alluauditoma Dajoz, 1980
- Anarmostes Pascoe, 1860
- Anisopaulax Reitter, 1877
- Anosyana Dajoz, 1980
- Antilissus Sharp, 1879
- Antillemonomma Freude, 1955
- Aplanetes Sharp, 1899
- Aprostoma Guérin-Méneville, 1839
- Ascomma Waterhouse, 1876
- Aspathines Champion, 1888
- Aspinatimonomma Freude, 1955
- Asprotera Pascoe, 1860
- Asynchita Sharp, 1894
- Atyscus Pascoe, 1863
- Aulonium Erichson, 1845
- Bhutania Dajoz, 1975
- Bitoma Herbst, 1793
- Bolcocius Dajoz, 1977
- Bulasconotus Slipinski & Lawrence, 1997
- Bulasconotus scaccarius Bernard & Gillett, 2020[7]
- Bupala Pascoe, 1863
- Caanthus Champion, 1894
- Cacotarphius Sharp, 1894
- Canaricerus Oromi & Garcia, 1987
- Caprodes Pascoe, 1863
- Carbothrus Aoki, 2012
- Catolaemus Sharp, 1894
- Cebia Pascoe, 1863
- Cerchanotus Erichson, 1845
- Chorasus Sharp, 1882
- Cicablabus Slipinski & Lawrence, 1997
- Ciconissus Broun, 1893
- Cleteomonomma Freude, 1957
- Cleteus Fairmaire, 1906
- Coeloderes Mulsant & Rey, 1859
- Colobicones Grouvelle, 1918
- Colobicus Latreille, 1807
- Colydium Fabricius, 1792
- Colydodes Motschulsky, 1855
- Corticus Latreille, 1829
- Coslonatus Li & Cai, 2021
- Cotulades Pascoe, 1860
- Coxelus Dejean, 1821
- Cretomysteria Deng, Ślipiński, Ren & Pang, 2017
- Curtemonomma
- Dechomus Jacquelin Du Val, 1859
- Denophloeus Stephan, 1989
- Denophoelus Stephan, 1989
- Diodesma Latreille, 1829
- Diplagia Reitter, 1882
- Diplotoma Erichson, 1845
- Ditomoidea Grouvelle, 1906
- Docalis Pascoe, 1860
- Doderonymus Binaghi, 1937
- Dryptops Broun, 1882
- Emilka Slipinski, 1982
- Enarsus Pascoe, 1866
- Endeitoma Sharp, 1894
- Endestes Pascoe, 1863
- Endocoxelus Waterhouse, 1876
- Endophloeus Dejean, 1834
- Enhypnon Carter, 1919
- Epistranodes Slipinski & Lawrence, 1997
- Epistranus Sharp, 1878
- Erylus Dajoz, 1969
- Eucicones Sharp, 1894
- Eudesma LeConte, 1863
- Faecula Slipinski & Lawrence, 1997
- Fenerivia Dajoz, 1980
- Franzorphius Schuh, 1998
- Gathocles Broun, 1893
- Gempylodes Pascoe, 1863
- Glenentela Broun, 1893
- Glyphocryptus Sharp, 1885
- Grandemonomma Freude, 1957
- Heterargus Sharp, 1886
- Holopleuridia Reitter, 1876
- Hyberis Pascoe, 1860
- Hybonotus Slipinski & Lawrence, 1997
- Hyboonotus Slipinski & Lawrence, 2016
- Hyporhagus Dejean, 1833
- Hyporhagus Thomson, 1860
- Hyporrhagus Thomson, 1860
- Hystricones Sharp, 1894
- Inscutomonomma Pic, 1951
- Insulcicleteus Freude, 1957
- Isotarphius Heller, 1916
- Kanantsia Dajoz, 1980
- Kraatziana Pic, 1934
- Labrotrichus Sharp, 1894
- Langelandia Aubé, 1842
- Lascobitoma Slipinski & Lawrence, 1997
- Lasconotus Erichson, 1845
- Lascotonus Grouvelle, 1895
- Lascotrichus Pope, 1961
- Lastrema Reitter, 1882
- Latometus Erichson, 1842
- Lemmis Pascoe, 1860
- Lepidosynchis Kolbe, 1897
- Linophloeus Dajoz, 1980
- Lobogestoria Reitter, 1878
- Lobomesa Slipinski & Lawrence, 1997
- Lyreus Aubé, 1861
- Madacones Dajoz, 1980
- Madadesia Dajoz, 1986
- Madenphloeus Dajoz, 1980
- Mamakius Pope, 1961
- Mauritianopidium Dajoz, 1977
- Mecedanum Erichson, 1845
- Megataphrus Casey, 1890
- Metopiestes
- Micromonomma Pic, 1933
- Microprius Fairmaire, 1868
- Microsicus Sharp, 1894
- Microvonus
- Monoedus Horn, 1882
- Monomma Klug, 1833
- Munaria Reitter, 1882
- Myrmecoxenus
- Namunaria Reitter, 1882
- Nematidium Erichson, 1845
- Neotrichus Sharp, 1886
- Niphopelta Reitter, 1882
- Norix Broun, 1893
- Noserinus Casey, 1907
- Nosoderma Guérin-Méneville, 1838
- Nosodomodes Reitter, 1922
- Notocoxelus Slipinski & Lawrence, 1997
- Notorthocerus Slipinski & Lawrence, 1999
- Opostirus Kirsch, 1865
- Orthocerodes Slipinski & Lawrence, 1999
- Orthocerus Latreille, 1796
- Orvoenia Dajoz, 1980
- Paha Dajoz, 1984
- Papuatrichus Aoki, 2013
- Paratarphius Dajoz, 1971
- Paryphus Erichson, 1845
- Paxillobitoma Lord & Ivie, 2016
- Phaennis Champion, 1894
- Pharax Pascoe, 1860
- Phellopsis LeConte, 1862
- Phloeodalis Erichson, 1845
- Phloeodes LeConte, 1862
- Phloeonemus Erichson, 1845
- Phloeopsidius Gebien, 1925
- Phormesa Pascoe, 1863
- Phorminx Carter & Zeck, 1937
- Phreatus Pascoe, 1863
- Plagiope Erichson, 1845
- Priolomopsis Dajoz, 1982
- Priolomus Erichson, 1845
- Pristoderes
- Pristoderus Hope, 1840
- Prostheca Wollaston, 1860
- Protarphius Broun, 1893
- Pseudaulonium Reitter, 1877
- Pseudendestes Lawrence, 1980
- Pseudocorticus Hinton, 1935
- Pseudotarphius Wollaston, 1873
- Pycnomerodes Broun, 1886
- Pycnomerus Erichson, 1842
- Rechodes Erichson, 1845
- Recyntus Broun, 1882
- Rhagodera Mannerheim, 1843
- Rhagoderidea Wickham, 1914
- Rhizonium Sharp, 1876
- Rhopalocerus Redtenbacher, 1842
- Rytinotus Broun, 1880
- Sallachius Pope, 1959
- Sasajia Masumoto & Akita, 2007
- Sassaka Pope, 1961
- Scoriaderma Fairmaire, 1894
- Sechellotoma Dajoz, 1980
- Semicoxelus Alekseev & Pankowski, 2020
- Sesaspis
- Sparactus Erichson, 1845
- Spinatimonomma Freude, 1955
- Spinhyporhagus Freude, 1984
- Sprecodes Pope, 1961
- Stenomonoedus Heinze, 1954
- Stephaniolus Ivie, Slipinsky & Wegrzynowicz, 2001
- Sympanotus Sharp, 1885
- Synagathis Carter & Zeck, 1937
- Synchita Hellwig, 1792
- Tarphiablabus Slipinski & Lawrence, 1997
- Tarphiomimus Wollaston, 1873
- Tarphiosoma Wollaston, 1862
- Tarphius Erichson, 1845
- Tentablabus Slipinski & Lawrence, 1997
- Todima Grouvelle, 1893
- Todimopsis Slipinski & Lawrence, 1997
- Trachypholis Erichson, 1845
- Trigonophloeus Dajoz, 1980
- Uloncrus
- Usechimorpha Blaisdell, 1929
- Usechus Motschulsky, 1845
- Verodes Casey
- Vitiacus Broun, 1893
- Xylolaemus Dejean, 1835
- Xylolaemus Redtenbacher, 1857
- Zanclea Pascoe, 1863
- Zopher Slipinski & Lawrence, 1999
- Zopherosis White, 1859
- Zopherus Gray, 1832
- Zopherus Laporte, 1840
古昆虫学
目前所知的瘤拟步行虫科化石纪录相当稀少，多数化石纪录来自波罗的海琥珀（始新世）[8][9][10]，最早的化石纪录为来自上白垩纪缅甸琥珀的 Paleoendeitoma antennata、Paleoendeitoma minuta（属于细坚虫亚科 Colydiinae）和 Cretomysteria burmanica （亚科未定论）[11]。
参考文献
- ^ Zopheridae Report. Integrated Taxonomic Information System. [2018-04-23]. （原始内容存档于2020-11-02）.
- ^ Browse Zopheridae. Catalogue of Life. [2018-04-23]. （原始内容存档于2019-02-07）.
- ^ Ślipiński, Stanisław Adam; Ślipiński, Stanisław Adam; Lawrence, John Francis; Lawrence, John Francis. Phylogeny and classification of Zopheridae sensu novo (Coleoptera: Tenebrionoidea) with a review of the genera of Zopherinae (excluding Monommatini). Muzeum i Instytut Zoologii PAN. 1999.
- ^ Mckenna, Duane D.; Wild, Alexander L.; Kanda, Kojun; Bellamy, Charles L.; Beutel, Rolf G.; Caterino, Michael S.; Farnum, Charles W.; Hawks, David C.; Ivie, Michael A. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology. 2015-10-01, 40 (4): 835–880 [2017-09-26]. ISSN 1365-3113. doi:10.1111/syen.12132. （原始内容存档于2017-08-23） （英语）.
- ^ Gunter, Nicole L.; Levkaničová, Zuzana; Weir, Tom H.; Ślipiński, Adam; Cameron, Stephen L.; Bocak, Ladislav. Towards a phylogeny of the Tenebrionoidea (Coleoptera). Molecular Phylogenetics and Evolution. 2014-10-01, 79: 305–312. ISSN 1055-7903. doi:10.1016/j.ympev.2014.05.028 （英语）.
- ^ Bocak, Ladislav; Barton, Christopher; Crampton-Platt, Alex; Chesters, Douglas; Ahrens, Dirk; Vogler, Alfried P. Building the Coleoptera tree-of-life for >8000 species: composition of public DNA data and fit with Linnaean classification. Systematic Entomology. 2014-01-01, 39 (1): 97–110 [2017-09-26]. ISSN 1365-3113. doi:10.1111/syen.12037. （原始内容存档于2017-09-27） （英语）.
- ^ Jared Bernard, Conrad P. D. T. Gillett. 2020. A New Species of cylindrical Bark Beetle (Zopheridae: Colydiinae) from Vanuatu establishes in Hawaii. Zootaxa. 4809(3); 593–599. DOI: doi.org/10.11646/zootaxa.4809.3.12
- ^ Alekseev, V.I. Lord, N.P.. A new species of Xylolaemus (Coleoptera: Zopheridae: Colydiinae) from Baltic amber. (PDF). Baltic journal of Coleopterology. 2014, 1 (14): 97–102 [2017-09-26]. （原始内容存档 (PDF)于2017-08-12）.
- ^ Alekseev, V.I. New cylindrical bark (Coleoptera: Zopheridae) from Baltic amber. (PDF). Baltic Journal of Coleopterology. 2015, 1 (15): 17–24 [2017-09-26]. （原始内容存档 (PDF)于2016-03-04）.
- ^ Alekseev, Vitalii I.; Bukejs, Andris. New Zopheridae (Coleoptera: Tenebrionoidea) from Baltic amber. Zootaxa. 2016-10-25, 4178 (3): 409–427 [2017-09-26]. ISSN 1175-5334. PMID 27811716. （原始内容存档于2019-01-29）.
- ^ Deng, Congshuang; Ślipiński, Adam; Ren, Dong; Pang, Hong. The first Mesozoic colydiid beetles (Coleoptera: Zopheridae: Colydiinae) from the Upper Cretaceous amber of Myanmar. Cretaceous Research. 2017-10-01, 78 (Supplement C): 71–77. doi:10.1016/j.cretres.2017.04.010.
