心網蟲屬
鏈狀心網蟲![]() | |
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科学分类 ![]() | |
界: | 动物界 Animalia |
总门: | 蜕皮动物总门 Ecdysozoa |
演化支: | 泛节肢动物 Panarthropoda |
並系群: | †葉足動物 Lobopodia |
纲: | †異蟲綱 Xenusia |
目: | †堅有爪目 Scleronychophora |
演化支: | †怪誕蟲類群 Hallucigeniids |
科: | †心網蟲科 Cardiodictyidae Hou & Bergstrom, 1995[3] |
属: | †心網蟲屬 Cardiodictyon Hou, Ramsköld, and Bergström, 1991[1] |
种: | 鏈狀心網蟲 C. catenulum
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二名法 | |
Cardiodictyon catenulum Hou, Ramsköld, and Bergström, 1991
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鏈狀心網蟲(學名:Cardiodictyon catenulum[1])是心網蟲科(學名:Cardiodictyidae[3])心網蟲屬(學名:Cardiodictyon[1])的唯一一個物種[1][4],也是模式種。此物種在中國南方發現的。[3][1]生存在中寒武紀。[1]此物種的化石標本非常普遍,總共發現多達兩百多個心網蟲的化石標本,而且大多發現的標本是扁的,而且是彎曲的,[4]再加上化石保存都很好。[5]心網蟲的頭部具有兩對的前附肢[3][6],頭裡還有可能類似大腦的器官[7][8][9][10][11],頭部區域是箭頭狀,前面的邊緣是圓形。其頭部由一對大骨片保護,頭部前緣有雙重結構[3],並有一對眼點[12]; 背板的形狀也被重新詮釋。[3]嘴巴和肛門位於身體前後的末端。[8]身體很細長還有一條條的環紋[13],23對邊邊很尖的六角形的骨片,還有23對腳,而腳上兩個爪子。[3][1]與怪誕蟲屬(Hallucigenia)其中的一種強壯怪誕蟲 (Hallucigenia fortis),因為既有身長的身軀[14][15][16],與心網蟲一樣都是在中國發現的。[1]與囉哩山蟲屬(Luolishania)、微網蟲屬(Microdictyon)、貧腿蟲屬(Paucipodia)[17]、爪網蟲屬 (Onychodictyon)[1]都是在中國早期發現的基群泛節肢生物。[6][18]
命名
心網蟲屬是在中國雲南省澂江市的澄江化石地發現的。[3][4][19]是源自希臘語kardia是指心, 和dictyon是網子的意思。catenula是源自拉丁語, 意指小鍊條或是鐵鍊,是指心網蟲身上的一排一排的骨板。[3]
描述
![](http://images.weserv.nl/?url=//upload.wikimedia.org/wikipedia/commons/thumb/9/98/20210926_Cardiodictyon_catenulum_diagrammatic_reconstruction.png/708px-20210926_Cardiodictyon_catenulum_diagrammatic_reconstruction.png)
![](http://images.weserv.nl/?url=//upload.wikimedia.org/wikipedia/commons/thumb/b/b7/Xenusion.png/199px-Xenusion.png)
頭部看起來像是橢圓形或是半圓形[4][20],頭部有原本被發現有三對細長的前附肢,其中兩對已經確定存在,第三對當時還不能確定[1],而之後的研究發現只有兩對前附肢。[6][21][20][22],位在頭部骨片的更前面。[20]頭部的一端被一對大骨片覆蓋[3][13][20][22][23],頭部有分節[6],沒有大骨片的頭部看起來就像末端[1],頭部和尾部的末端逐漸變得細。[3]在頭部上還有細小的眼點。[3][4][12][20][24]身體細長[3][4],大約身長3公分、1公分寬[4],身體沒有分節[6],還有25對附肢(頭部的兩對附肢加上身體的二十三對用來行走的腳)[25][26][27],共有23塊的網狀六角形骨片[3][13][25][26][28][註 1],每個骨片與骨片的間隔總是有規律的,最後一塊的骨片在後面就是身體的尾部,而且尾部是鈍的。[3]原本以為身上是沒有環紋[1],後來證明有一條條的環紋。[13]發現有一從頭部延伸身體的末端條縱向的暗黑色寬帶,推測可能是腸子。[3][4][13][29]整個身體的骨板形狀幾乎沒什麼變化,每隔一對骨板,也就有一對腳。[3]這一些腳很長,大約是身體寬度的三倍。[1]當初而底部的爪子保存的不完整[1],因而誤以為有四、五個爪子[1][3],而之後發現其實只有兩個爪子。[25]
![](http://images.weserv.nl/?url=//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Anomalocaris_ecological.png/209px-Anomalocaris_ecological.png)
生活
對心網蟲的生活幾乎一無所知,有一些甚至是以對其他寒武紀葉足動物認知而成的。心網蟲是底棲型的物種,因為具有用來行走的腳,發現的腸子可能代表心網蟲是肉食性或是雜食性,且頭上兩對的附肢可能與放射齒目一樣,用來抓捕獵物。[4]
資料來源
- ^ 1.00 1.01 1.02 1.03 1.04 1.05 1.06 1.07 1.08 1.09 1.10 1.11 1.12 1.13 1.14 1.15 XIANGUANG, HOU; RAMSKÖLD, LARS; BERGSTRÖM, JAN. Composition and preservation of the Chengjiang fauna –a Lower Cambrian soft‐bodied biota. Zoologica Scripta. 1991-10, 20 (4). ISSN 0300-3256. doi:10.1111/j.1463-6409.1991.tb00303.x.
- ^ Ou, Qiang; Mayer, Georg. A Cambrian unarmoured lobopodian, †Lenisambulatrix humboldti gen. et sp. nov., compared with new material of †Diania cactiformis. Scientific Reports. 2018-09-20, 8 (1). ISSN 2045-2322. doi:10.1038/s41598-018-31499-y.
- ^ 3.00 3.01 3.02 3.03 3.04 3.05 3.06 3.07 3.08 3.09 3.10 3.11 3.12 3.13 3.14 3.15 3.16 3.17 HOU, XIANGUANG; BERGSTRÖM, JAN. Cambrian lobopodians-ancestors of extant onychophorans? (PDF). Zoological Journal of the Linnean Society. 1995-05, 114 (1). ISSN 0024-4082. doi:10.1111/j.1096-3642.1995.tb00110.x.
- ^ 4.0 4.1 4.2 4.3 4.4 4.5 4.6 4.7 4.8 Hou, X., Aldridge, R. J., Bergström, J., Siveter, D. J., Siveter, D. J., & Xiang-Hong, F. (2004). The Cambrian fossils of Chengjiang, China: the flowering of early animal life (Vol. 233, pp. 1-233). Oxford: Blackwell.
- ^ Han, Jian; Shu, Degan; Zhang, Zhifei; Liu, Jianni; Zhang, Xingliang; Yao, Yang. Preliminary notes on soft-bodied fossil concentrations from the Early Cambrian Chengjiang deposits. Chinese Science Bulletin. 2006-10, 51 (20). ISSN 1001-6538. doi:10.1007/s11434-005-2151-0.
- ^ 6.0 6.1 6.2 6.3 6.4 Chen, Jun-Yuan. The sudden appearance of diverse animal body plansduring the Cambrian explosion. The International Journal of Developmental Biology. 2009, 53 (5-6). ISSN 0214-6282. doi:10.1387/ijdb.072513cj.
- ^ Smith, Frank W.; Game, Mandy; Mapalo, Marc A.; Chavarria, Raul A.; Harrison, Taylor R.; Janssen, Ralf. Developmental and genomic insight into the origin of the tardigrade body plan. Evolution & Development. 2023-09-18. ISSN 1520-541X. doi:10.1111/ede.12457.
- ^ 8.0 8.1 Strausfeld, Nicholas J.; Hou, Xianguang; Sayre, Marcel E.; Hirth, Frank. The lower Cambrian lobopodian Cardiodictyon resolves the origin of euarthropod brains. Science. 2022-11-25, 378 (6622). ISSN 0036-8075. doi:10.1126/science.abn6264.
- ^ Budd, Graham E.; Mayer, Georg; Janssen, Ralf; Eriksson, B. Joakim. Comment on “The lower Cambrian lobopodian Cardiodictyonresolves the origin of euarthropod brains”. Science. 2023-06-30, 380 (6652). ISSN 0036-8075. doi:10.1126/science.adg1412.
- ^ Strausfeld, Nicholas J.; Hou, Xianguang; Sayre, Marcel E.; Hirth, Frank. Response to Comment on “The lower Cambrian lobopodian Cardiodictyon resolves the origin of euarthropod brains”. Science. 2023-06-30, 380 (6652). ISSN 0036-8075. doi:10.1126/science.adg6051.
- ^ Briggs, Derek E. G.; Parry, Luke A. Putting heads together. Science. 2022-11-25, 378 (6622). ISSN 0036-8075. doi:10.1126/science.add7372.
- ^ 12.0 12.1 Ma, Xiaoya; Hou, Xianguang; Aldridge, Richard J.; Siveter, David J.; Siveter, Derek J.; Gabbott, Sarah E.; Purnell, Mark A.; Parker, Andrew R.; Edgecombe, Gregory D. Morphology of Cambrian lobopodian eyes from the Chengjiang Lagerstätte and their evolutionary significance. Arthropod Structure & Development. 2012-09, 41 (5). ISSN 1467-8039. doi:10.1016/j.asd.2012.03.002.
- ^ 13.0 13.1 13.2 13.3 13.4 Waloszek, Dieter; Maas, Andreas; Chen, Junyuan; Stein, Martin. Evolution of cephalic feeding structures and the phylogeny of Arthropoda. Palaeogeography, Palaeoclimatology, Palaeoecology. 2007-10, 254 (1-2). ISSN 0031-0182. doi:10.1016/j.palaeo.2007.03.027.
- ^ Smith, Martin R.; Ortega-Hernández, Javier. Hallucigenia’s onychophoran-like claws and the case for Tactopoda. Nature. 2014-08-17, 514 (7522). ISSN 0028-0836. doi:10.1038/nature13576.
- ^ Smith, Martin R.; Caron, Jean-Bernard. Hallucigenia’s head and the pharyngeal armature of early ecdysozoans. Nature. 2015-06-24, 523 (7558). ISSN 0028-0836. doi:10.1038/nature14573.
- ^ Yang, Jie; Ortega-Hernández, Javier; Gerber, Sylvain; Butterfield, Nicholas J.; Hou, Jin-bo; Lan, Tian; Zhang, Xi-guang. A superarmored lobopodian from the Cambrian of China and early disparity in the evolution of Onychophora. Proceedings of the National Academy of Sciences. 2015-06-29, 112 (28). ISSN 0027-8424. doi:10.1073/pnas.1505596112.
- ^ Jun-yuan, Chen; Gui-qing, Zhou; Ramsköld, Lars. A new Early Cambrian onychophoran-like animal, Paucipodia gen. nov., from the Chengjiang fauna, China. Transactions of the Royal Society of Edinburgh: Earth Sciences. 1994, 85 (4). ISSN 0263-5933. doi:10.1017/s0263593300002042.
- ^ Vannier, Jean; Liu, Jianni; Lerosey-Aubril, Rudy; Vinther, Jakob; Daley, Allison C. Sophisticated digestive systems in early arthropods. Nature Communications. 2014-05-02, 5 (1). ISSN 2041-1723. doi:10.1038/ncomms4641.
- ^ Maas, Andreas; Mayer, Georg; Kristensen, Reinhardt M.; Waloszek, Dieter. A Cambrian micro-lobopodian and the evolution of arthropod locomotion and reproduction. Chinese Science Bulletin. 2007-12, 52 (24). ISSN 1001-6538. doi:10.1007/s11434-007-0515-3.
- ^ 20.0 20.1 20.2 20.3 20.4 Ma, Xiaoya; Hou, Xianguang; Bergström, Jan. Morphology of Luolishania longicruris (Lower Cambrian, Chengjiang Lagerstätte, SW China) and the phylogenetic relationships within lobopodians. Arthropod Structure & Development. 2009-07, 38 (4). ISSN 1467-8039. doi:10.1016/j.asd.2009.03.001.
- ^ Dzik, J. (2011). The xenusian-to-anomalocaridid transition within the lobopodians. (页面存档备份,存于互联网档案馆) Boll Della Soc Paleontol Ital, 50, 65-74.
- ^ 22.0 22.1 Liu, JianNi; Shu, DeGan; Han, Jian; Zhang, ZhiFei. Comparative study of Cambrian lobopods Miraluolishania and Luolishania. Chinese Science Bulletin. 2008-01, 53 (1). ISSN 1001-6538. doi:10.1007/s11434-007-0428-1.
- ^ Bergström, J.a.n.; Hou, Xian-Guang. Cambrian Onychophora or Xenusians. Zoologischer Anzeiger - A Journal of Comparative Zoology. 2001-01, 240 (3-4). ISSN 0044-5231. doi:10.1078/0044-5231-00031.
- ^ Liu, Jianni; Dunlop, Jason A. Cambrian lobopodians: A review of recent progress in our understanding of their morphology and evolution. Palaeogeography, Palaeoclimatology, Palaeoecology. 2014-03, 398. ISSN 0031-0182. doi:10.1016/j.palaeo.2013.06.008.
- ^ 25.0 25.1 25.2 Liu, Jianni; Shu, Degan; Han, Jian; Zhang, Zhifei; Zhang, Xingliang. Origin, diversification, and relationships of Cambrian lobopods. Gondwana Research. 2008-08, 14 (1-2). ISSN 1342-937X. doi:10.1016/j.gr.2007.10.001.
- ^ 26.0 26.1 Bergström, J.a.n.; Hou, Xian-Guang. Cambrian Onychophora or Xenusians. Zoologischer Anzeiger - A Journal of Comparative Zoology. 2001-01, 240 (3-4). ISSN 0044-5231. doi:10.1078/0044-5231-00031.
- ^ Siveter, Derek J.; Briggs, Derek E. G.; Siveter, David J.; Sutton, Mark D.; Legg, David. A three-dimensionally preserved lobopodian from the Herefordshire (Silurian) Lagerstätte, UK. Royal Society Open Science. 2018-08, 5 (8). ISSN 2054-5703. doi:10.1098/rsos.172101.
- ^ Smith, Martin Ross; Dhungana, Alavya. Discussion on Tabelliscolex (Cricocosmiidae: Palaeoscolecidomorpha) from the early Cambrian Chengjiang Biota and the evolution of seriation in Ecdysozoa’ by Shiet al.2021 (JGS, jgs2021-060). Journal of the Geological Society. 2022-01-28, 179 (3). ISSN 0016-7649. doi:10.1144/jgs2021-111.
- ^ Hou, Xian‐Guang; Ma, Xiao‐Ya; Zhao, Jie; Bergström, Jan. The lobopodian Paucipodia inermis from the Lower Cambrian Chengjiang fauna, Yunnan, China. Lethaia. 2004-09, 37 (3). ISSN 0024-1164. doi:10.1080/00241160410006555.
- ^ 通常發現心網蟲化石標本都是發現側面的,所以骨片看起來類似三角形;如果以背側骨片看,那看到的是類似六角形。
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